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Police found the victims just after 1 a.m. in the northbound lanes of I-95 near Exit 2 in Upper Chichester Township.
When police arrived at the scene, they discovered a 41-year-old man from Brookhaven unresponsive inside the car. He was pronounced dead at the scene.
Police say another man, a 42-year-old passenger, was also inside the car suffering from multiple gunshot wounds.
He was taken to Crozer Medical Center. His condition is unknown at this time.
The northbound lanes of I-95 were shut down between Exits 2 and 3 from 1 a.m. until 10 a.m. while Pennsylvania State Police investigated.
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All lanes have since re-opened.
This is an ongoing investigation with state police.