Philly Zoo mourns Makini, a 17-year-old African lion

PHILADELPHIA (WPVI) -- The Philadelphia Zoo is mourning the loss of one of its beloved lions.

Makini, a 17-and-a-half-year-old African lion, was humanely euthanized after severe arthritis left him weak and struggling to walk.

Makini spent most of his life at the zoo and fathered four cubs.

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The zoo says it's now working with the Association of Zoos and Aquariums on the future of lions there.